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Besides those already mentioned, there are some other meanings of the skulls tattoos that have their roots in some ancient beliefs and practices.
Skulls as trophies - it was a common practice to keep the skulls of the killed enemies and to display them as trophies (eg.: to decorate with them the city walls and towers, to keep the skulls on the palisades as a warning sign, to display them on pikes at the Traitor's Gate, etc) or even to use them as wine-cups. Skulls-tattoos as trophies are a very common theme when it comes to the gang tattoos, prison tattoos, etc. For example, if we consider the russian criminal tattoos, a skull tattoo signifies murder (a significant one). When a tattoo design combines skulls with military insignia (especially with the epaulets tattooed on the shoulders), it usually designates a murderer.

The epaulets are distinctive decorations for the inmates who are members of the "Vory V Zakone" (aprox "Thieves in law"). An inmate who has this kind of tattoo is a person with a high position in the prison hierarchy. The presence of the skull designates the wearer as a murderer. Read more about this topic on this blog: russiancriminaltattoo.blogspot.com

A russian criminal tattoo. The presence of the epaulets indicates that the wearer has a high rank in the prison hierarchy. The presence of the skull designates the wearer as a murderer. The spider is a symbol with multiple meanings; it may symbolize racism, doing time in prison, drug addicted, etc. Read more about this topic on this blog: russiancriminaltattoo.blogspot.com
In the Mexican tattoo-culture, skulls and skeletons are linked with the celebration of the Day of the Dead. The most common themes and representations are: the sugar-skull (brightly colored and highly decorated), the partying skeletons (because El Dia de los Muertos is not a time for sadness, but for joy and celebration) and the in-loving-memory-tattoos (these tattoos include attributes of the passed person in the skull's design: her favourite flowers, his / her weaopns, facial features, etc).

In-loving-memory-tattoo. Some of the passed person's features are still present (the moustache and the hair)

Typical Mexican tattoo: a fancy dressed skeleton playing a violin

For the Mexicans, El Dia de los Muertos is not a mournful day, but a day of family gatherings and holiday cheer.
Alchemy and Magic - the skull is an ancient symbol and it has several embodiments of the alchemical �union of opposites�, an ancient doctrine which accepts a syncretism of conflicting views, a harmony as opposed to a never-ending conflict between Good and Evil.
Wisdom - sometimes, a skull tattoo is a symbol of the mature realization of a continuity of life into death, and the ability to accept death. This is very important because it is only when a person can live with the presence of death that he can fully learn to appreciate life.
Related posts:
Various elements which can occur in a Death tattoo: skulls and skeletons (1)
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Many people choose to get a skull tattoo because they want to show their affiliation to a group or to show their beliefs and their devotion to them.
The famous Jolly Roger (the black flag with a white skull and two human bones crossed under the skull) once used by pirates, was both an intimidating signal and a symbol of the total disregard for the authorities, as well as a symbol of the absolute freedom. This symbol (which is still preserving its original meaning) symbol was integrated into various systems of symbols belonging to different subculture or/and counterculture groups (like the outlaw motorcycle clubs whose members often get Jolly Roger tattoos to show their lawlessness). Sometimes, the anarchists use the "Jolly Roger" as a symbol of rebellion against the organized authority.
The Dead Man's Head consists usually of the human skull with or without the mandible and often includes two crossed long-bones (the femurs). Unlike the Jolly Roger design, the Dead Man's Head design features a skull with the crossbones placed behind the skull, instead of beneath it. The Dead Man's Head is an ancient symbol and has had different meanings over time; in the Elizabethan England, for instance, The Death's-Head Skull (usually a depiction without the lower jawbone), was emblematic of bawds, rakes, sexual adventurers and prostitutes; the term Deaths-Head was actually parlance for these rakes, and most of them wore half-skull rings to advertise their station, either professionally or otherwise. The original rings were wide silver objects, with a half-skull decoration not much wider than the rest of the band; this allowed it to be rotated around the finger to hide the skull in polite company, and to reposition it in the presence of likely conquests.
The Totenkopf was used by the german Schutzstaffel (SS) as the unit's insignia. As a consequence of that, this particular design of the Dead Man's Head has become closely associated with various movements like the "white nationalist" movements, the "white power skinhead" movements and the "Neo-Nazi" movement.

a Totenkopf tattoo - a tattoo design closely associated with the Neo-Nazi's ideology
the 3rd SS Division's Totenkopf insignia

Totenkopf design; the draw also contain a variant of so-called "Cross Pattee (English: Cross Patty; German: Tatzenkreuz)"; the Cross Pattee is often mistakenly identified with the gothic crosses, but actually it is an ancient symbol which was largely used by the Teutonic Knights and sometimes by the Templiers. The Cross Pattee design was incorporated in the design of the "Iron Cross", a german decoration for military valor established by the King Frederick William III of Prussia in 1813. The Iron Cross remained in use, in various forms, by Prussia and later Germany until 1945.
Variations on the Skull and Crossbones are used by various military units as a symbol of strength, courage and bravure, as well as a symbol of the disregard for one's life. On wikipedia you can find a list of the Skull and crossbones military insignias.

The BOPE emblem, a skull impaled on a sword, backed by two gold pistols sends out a strong, unambiguous message. The emblem symbolises armed combat, war and death. The BOPE�s emblem, a skull with a crisscross of pistols, is believed to possess mystical powers of protection. BOPE = Special Policial Operations Battalion, a special forces unit of the Military Police of Rio de Janeiro State, Brazil. If you want to know more about BOPE you can visit their blog or you can watch these two movies: Elite Squad and Elite Squad: The Enemy Within
Related posts:
Various elements which can occur in a Death tattoo: skulls and skeletons (1)
Death tattoo - various elements which can occur in a Death tattoo
Death tattoos: the main depictions of the Grim Reaper
Death Tattoos / Grim Reaper Tattoos: themes and meanings
Death tattoo - various elements which can occur in a Death tattoo - animals
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Human skulls and skeletons are one of the most common elements in the Grim Reaper tattoos, due to the fact that skeletons and skulls have long been associated with death and decay (sometimes Death himself is portrayed as a skeleton).
Skull tattoos do represent death, but they usually represent the fleetingness of life. Various Latin maxims and aphorisms: Memento Mori (Remember your mortality, Remember you must die or Remember you will die), Sic Transit Gloria Mundi (Thus passes the glory of the world), Carpe Diem (Seize the day), Ubi sunt...? (Where are...?) or Ubi nunc...? (Where now...?) - both of them are shortened variants of the Ubi sunt qui ante nos fuerunt? Where are those who were before us?) are very often included in the skull tattoo designs; their role is to remind us that Death comes to us all, and for this reason is important to enjoy every moment of life.

Seize the day or die regretting - a tattoo design which contains flowers (which symbolize brevity and the ephemeral nature of life) and a skull
As a symbol of the certainty of death, skulls are one of the most common symbols used in the Vanitas art

vanitas tattoo containing a skull (a symbol of dead and mortality), several roses (symbols of the ephemeral nature of life), a candle (an object closely related with the cult of the dead) and a watch (a symbol of the inexorable flow of time)
Despite the fact that the skull is often a symbol of mortality, a design which depicts a skull backed by a pair of wings (usually a pair of angelic wings, but this is not mandatory) is rather a positive design with various meanings: resurrection; death is the road to a new form of existence; the death frees one of the burdens of life; the progression of the human soul through the afterlife. Anyway, the winged skull is not always a positive design, having also many negative connotations: danger, death (death which can reach everyone everywhere), lawlessness (very common in the bikers-gang tattoos)
The skull is also a symbol of the transitory nature of life and death, and for this reason some designers choose to include a skull into more complex designs which represent the infinite cycle of the life and death.
Related posts:
Death tattoo - various elements which can occur in a Death tattoo
Death tattoos: the main depictions of the Grim Reaper
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The most common elements in the Death tattoos are, without any doubt, the weapons (for the obvious reason that they are used for killing people).
The scythe is, by far, the most popular choice when it comes to a Grim Reaper tattoo. The scythe can symbolize the cutting of negative energies or influences. Also, as a harvesting tool, it can symbolize hope and prosperity. It can represent the moment of death, when a scythe is said to cut a silver thread. The scythe has a long handle, indicating that it can reach anywhere.

Death tattoo: a skeleton praying to Death. If you like this tattoo design you can find it in a larger format on my web site, www.seiza.ro
However, when the Death is portrayed as the Death Angel, the scythe is usually (but not necessarily) replaced by a sword. The sword can be a short, simple, sword (similar to the roman Gladius) or a highly-decorated sword.
Without being very popular, many other medieval weapons can be seen in the Death tattoos: spears, lances and javelins (these weapons can be seen in the Angel of Death tattoos or in the tattoos which depict Death as a medieval knight), knives and daggers (very common in the Skulls tattoo designs), bows and arrows, etc.
Increasingly popular are the fire-guns: usually handguns - derringers, old-fashioned revolvers (very common in the Mexican Mafia tattoos and in the Skulls tattoos), semi-automatic pistols and machine pistols (very common in the gang / prison tattoos) but also various long guns (like shotguns or sniper rifles).

Gangsta tattoo: Grim Reaper pointing a revolver. This tattoo was drawn by Peter Coinin Jr., a tattoo artist who works at the Beauty Mark Tattoo, 3356 East Main Street Waterbury, Connecticut. If you like this tattoo and you want to buy it you can send an email at this mail-address: beautymarktattoo@yahoo.com or you can visit their web-page

skull tattoo covering a man's chest: a skull and two crossing revolvers surrounded by several gunshot wounds
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